Jack Hall, paleoclimatologist, must make a daring trek from Washington, D.C. to New York City to reach his son, trapped in the cross-hairs of a sudden international storm which plunges the planet into a new Ice Age.

Manny, Sid and Diego discover that the ice age is coming to an end, and join everybody for a journey to higher ground. On the trip, they discover that Manny is not in fact the last of the woolly mammoths.

When Scrat’s acorn sends an asteroid to Earth, the Herd must find a way to stop the asteroid from hitting Earth with the help of a returning friend.