Inside of his book, adventurous Harold can make anything come to life simply by drawing it. After he grows up and draws himself off the book’s pages and into the physical world, Harold finds he has a lot to learn about real life.

It’s a jungle out there for Blu, Jewel, and their three kids after they’re hurtled from Rio de Janeiro to the wilds of the Amazon. As Blu tries to fit in, he goes beak-to-beak with the vengeful Nigel and meets his father-in-law.

Manny, Sid and Diego discover that the ice age is coming to an end, and join everybody for a journey to higher ground. On the trip, they discover that Manny is not in fact the last of the woolly mammoths.